［1］YANG K, JEONG N, MOON J K, et al. Genetic analysis of genes controlling natural variation of seed coat and flower colors in soybean[J]. Journal of Heredity, 2010, 101(6): 757-768.[2]BALDONI E, GENGA A, COMINELLI E. Plant MYB transcription factors: Their role in drought response mechanisms[J]. International Journal of Molecular Sciences, 2015, 16(7): 15811-15851.[3]李海芬, 邱金梅, 陈小平, 等. 花生花青素合成相关基因的表达与种皮颜色关系[J]. 中国油料作物学报, 2017, 39(5): 600-605. (LI H F, QIU J M, CHEN X P. Differential expression of genes associated with anthocyanin synthesis in peanut cultivars with different testa color[J]. Chinese Journal of Oil Crop Sciences, 2017, 39(5): 600-605.)[4]FUJIWARA Y, KONO M, ITO A, et al. Anthocyanins in perilla plants and dried leaves[J]. Phytochemistry, 2018, 147: 158-166.[5]DONG T T, HAN R P, YU J W, et al. Anthocyanins accumu-lation and molecular analysis of correlated genes by metabolome and transcriptome in green and purple asparaguses (Asparagus oicinalis, L.)[J]. Food Chemistry, 2019, 271: 18-28.[6]ZHENG T, TAN W R, YANG H, et al. Regulation of anthocyanin accumulation via MYB75/HAT1/TPL-mediated transcriptional repression[J]. PLoS Genetics, 2019, 15(3): e1007993.[7]NESI N, JOND C, DEBEAUJON I, et al. The Arabidopsis TT2 gene encodes an R2R3 MYB domainprotein that acts as a key determinant for proanthocyanidin accumulation in developing seed[J]. Plant Cell, 2001, 13(9): 2099-2114.[8]TODDJ J, VODKIN L O. Duplications that suppress anddeletions that restore expression from a CHS multigene family[J]. Plant Cell, 1996, 8: 687-699.[9]ZABALA G, VODKIN L. Cloning of the pleiotropic T locus in soybean and two recessive alleles that differentially affect structure and expression of the encoded flavonoid 39 hydroxylase[J]. Genetics, 2003, 163: 295-309.[10]GRACIA Z, Vodkin L O. Methylation affects transposition and splicing of a large CACTA transposon from a MYB transcription factor regulating anthocyanin synthase genes in soybean seed coats[J]. PLoS One, 2014, 9(11): e111959. [11]WANG H Y, WANG H L, SHAO H B, et al. Recent advances in utilizing transcription factors to improve plant abiotic stress tolerance by transgenic technology[J]. Frontiers in Plant Science, 2016, 7: 67.[12]BOREVITZ J O, XIA Y, BLOUNT J, et al. Activation agging identifies a conserved MYB regulator of phenylpropanoid biosynthesis[J]. Plant Cell, 2000, 12: 2383-2393.[13]ZULUAGA D L, GONZALI S, LORETI E, et al. Arabidopsis thaliana MYB75/PAP1 transcription factor induces anthocyanin production in transgenic tomato plants[J]. Functional Plant Biology, 2008, 35(7): 606-618.[14]KIFERLE C, FANTINI E, BASSOLINO L, et al. Tomato R2R3-MYB proteins SlANT1 and SlAN2: Same protein activity, different roles[J]. PLoS One, 2015, 10(8): e0136365.[15]TOHGE T, NISHIYAMA Y, HIRAI M Y, et al. Functional genomics by integrated analysis of metabolome and transcriptome of Arabidopsis plants over-expressing an MYB transcription factor[J]. The Plant Journal, 2005, 42(2): 218-235.[16]DUBOS C, STRACKE R, GROTEWOLD E, et al. MYB trans-cription factors in Arabidopsis[J]. Trends Plant Science, 2010, 15: 573-581.[17]KATIYAR A, SMITA S, LENKA S K. Genome-wide classification and expression analysis of MYB transcription factor families in rice and Arabidopsis[J]. BMC Genomics, 2012, 13: 544-563.[18]JAVIER P A, DEBABROTA G, UDO W, et al. The regulatory cl locus of Zea mays encodes a protein with homology to myb proto-oncogene products and with structural similarities to transcriptional activators[J]. The EMBO Journal, 1987, 6(12): 3553-3558.[19]蒋文慧. 小麦中调控胚芽鞘与籽粒花青素合成相关基因的筛选及其功能验证[D]. 杨凌：西北农林科技大学, 2018: 1-52. (JIANG W H. Screening and functional identification of genes related to coleoptile and grain anthocyanin synthesis in wheat [D]. Yangling: Northwest Agriculture and Forestry University, 2018: 1-52.)[20]PENG X X, BAI N N, WANG H H. Isolation and expression profiles of cadmium stress-responsive rice WRKY15 transcription factor gene[J]. Chinese Journal of Rice Science, 2018, 32(2): 103-110.[21]LIU J Y, OSBOURN A, MA P D. MYB transcription factors as regulators of phenylpropaniod metabolism in plants[J]. Molecular Plant, 2015, 8: 689-708.[22]SONG J, LIU Z, HONG H, et al. Identification and validation of loci governing seed coat color by combining association mapping and bulk segregation analysis in soybean[J]. PLoS One, 2016, 12, 11(7): e0159064.[23]YAN F, DI S K, RYOJI T. CACTA-superfamily transposable element is inserted in MYB transcription factor gene of soybean line producing variegated seeds[J]. Genetics, 2015, 58(8): 365-374.[24]宋健. 大豆种皮色相关基因的图位克隆及功能解析[D]. 北京: 中国农业科学院, 2019: 82-85. (SONG J. Map based cloning and functional analysis of soybean seed coat color related genes[D]. Beijing: Chinese Academy of Agricultural Sciences, 2019: 82-85.)[25]宋健, 郭勇, 于丽杰, 等. 大豆种皮色相关基因研究进展[J]. 遗传, 2012, 34(6): 687-694. (SONG J, GUO Y, YU L J, et al. Research progress of soybean seed coat color related genes[J]. Genetics, 2012, 34 (6): 687-694.)[26]ZHAO Y, MA J, LI M, et al. Whole-genome resequencing-based QTL-seq identifiedAhTc1 gene encoding a R2R3-MYB transcription factor controlling peanut purple testa colour[J]. Plant Biotechnology Journal, 2020, 18(1): 96-105.[27]王关林.植物基因工程原理与技术[M]. 北京： 科学出版社, 1998： 86-90.(WANG G L.Principle and technology of plant genetic engineering［M］.Beijing: Science Press, 1998: 86-90.)[28]SERRANO M, KANEHARA K, TORRES M, et al. Repression of sucrose/ultraviolet B light-induced flavonoid accumulation in microbe-associated molecular pattern-triggered immunity in Arabidopsis[J]. Plant Physiophgy, 2012, 158(1): 408-422.[29] 张芷睿, 张耀华, 王秋实, 等. 大豆实时荧光定量PCR内参基因的筛选与验证[J].植物生理学报, 2020, 56(9): 1963-1973. (ZHANG Z R, ZHANG Y H, WANG Q S, et al. Screening and validation of internal reference genes in soybean real-time fluorescence quantitative PCR[J]. Journal of Plant Physiology, 2020, 56 (9): 1963-1973.)[30]蒋彦. 基础生物信息学及应用[M]. 北京: 清华大学出版社, 2003. (JIANG Y. Basic bioinformatics and application[M]. Beijing: Tsinghua University Press, 2003.)[31]LI Y, CHEN Q, XIE X, et al. Integrated metabolomics andtranscriptomics analyses reveal the molecular mechanisms underlying the accumulation of anthocyanins and other flavonoids in cowpea pod (Vigna unguiculata L.)[J]. Journal of Agricultural and Food Chemistry, 2020, 68(34): 9260-9275.[32]刘恺媛, 王茂良, 辛海波, 等.植物花青素合成与调控研究进展[J].中国农学通报,2021,37(14):41-51. (LIU K Y, WANG M L, XIN H B, et al. Anthocyanin biosynthesis and regulate mechanisms in plants: A review[J]. Chinese Agricultural Science Bulletin, 2021, 37(14): 41-51.)[33]TAMAGNONE L, MERIDA A, PARR A, et al. The AmMYB308 and AmMYB330 transcription factors from Antirrhinum regulate phenylpropanoid and lignin biosynthesis in transgenic tobacco[J]. Plant Cell, 1998, 10: 135-154.[34]吴嘉诚, 王桂青, 曾黎辉. 中国水仙R2R3-MYB基因NtMYB5的克隆和功能研究[J]. 园艺学报, 2018, 45(7): 1327-1337. (WU J C, LIU G Q, ZENG L H, Cloning and functional analysis of R2R3-MYB gene NtMYB5 in Narcissus tazetta var. chinensis[J]. Horticultural Plant Journal, 2018, 45(7): 1327-1337.)[35]ZHOU M L, ZHANG K X, SUN Z M, et al. LNK1 and LNK2 corepressors interact with the MYB3 transcription factor in phenylpropanoid biosynthesis[J]. Plant Physiology, 2017, 174(3): 1348-1358.[36]YANG L, ZHAO X, RAN L Y, et al. PtoMYB156 is involved in negative regulation of phenylpropanoid metabolism and secondary cell wall biosynthesis during wood formation in poplar[J]. Scientific Reports, 2017, 7: 41209.[37]XU Z S, FENG K, QUE F, et al. A MYB transcription factor DcMYB6 is involved in regulating anthocyanin biosynthesis in purple carrot taproots[J]. Scientific Reports, 2017(7): 45324.[38]李军, 赵爱春, 余茂德. 桑树花青素合成相关MYB类转录因子的鉴定与功能分析[J]. 西北植物学报, 2016, 36(6): 1110-1116. (LI J, ZHAO A C, YU M D. Identification and function analysis of anthocyanin biosynthesis related MYB genes in mulberry[J]. Acta Botanica Boreali Occidentalia Sinica, 2016, 36(6): 1110-1116.)[39]AKADA S, DUBE S K. Organization of chalcone synthase gene clusters and characterization of a new member of the family[J]. Plant Molecular Biology, 1995, 29: 189-199. [40]CHO Y B, JONES S I, VODKIN L O. Nonallelic homologous recombination events responsible for copy number variation within an RNA silencing locus[J]. Plant Direct, 2019, 3: e00162.[41]LIU Y C, DU H L, LI P C, et al. Pan-genome of wild and cultivated soybeans[J]. Cell, 2020, 182: 162-176.[42]GUO Y, QIU L J. Allele-specific marker development and selec-tion efficiencies for both flavonoid 3’-hydroxylase and flavonoid 3’,5’-hydroxylase genes in soybean subgenus soja[J]. Theoretical and Applied Genetics, 2013, 126: 1445-1455. [43]YAN F, DI S K, TAKAHASHI R. CACTA-superfamily trans-posable element is inserted in MYB transcription factor gene of soybean line producing variegated seeds[J]. Genetics, 2015, 58: 365-374. [44]ZABALA G, VODKIN L O. Methylation affects transposition and splicing of a large CACTA transposon from a MYB transcription factor regulating anthocyanin synthase genes in soybean seed coats[J]. PLoS One, 2014, 9: e111959. [45]徐金瑞, 张名位, 刘兴华, 等. 黑大豆种质抗氧化能力及其与总酚和花色苷含量的关系[J]. 中国农业科学, 2006, 39: 1545-1552. (XU J R, ZHANG M W, LIU X H, et al. Correlation between antioxidation, and content of total phenolics and anthocyanin in black soybean accessions[J]. Scientia Agricaltura Sinica, 2006, 39: 1545-1552.)[46]YOSHIDA K. Structural analysis and measurement of anthocyanins from colored seed coats of Vigna, Phaseolus, and Glycine legumes[J]. Bioscience, Biotechnology, and Biochemistry, 1996, 60: 589-593.